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Only after several inadequate breaths under such conditions,

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Only after several inadequate breaths under such conditions,resulting in the considerable elevation of carbon dioxide and reduction of oxygen in the blood,is breathing effort adjusted.谓语动词为什么是is不是are呢?这句话是后面的is breathing effort adjusted是什么意思
Only after several inadequate breaths under such conditions,
Breathing effort is adjusted only after...,resulting in ...是不是该这样理解?